Why do major political parties die? The shelf life of minor parties in democracies tends to be short, but major parties tend to be highly durable. The Democratic Party of the United States and the Conservative Party of the United Kingdom have been going strong for two centuries. Major parties perpetuate themselves by maintaining a consistent ideology on major national issues, even at the cost of periodic defeats at the polls. In American politics, ideological polarization maintains the vitality of the two major parties and renders them almost immune to threats from new parties, even as it impedes consensus and compromise on public issues.Spectacular instances of sudden death in major parties have nevertheless occurred in the United States, the United Kingdom, Canada, and Italy, and they all exhibit similar characteristics. The fatal event-which author Charles S. Mack calls "disalignment"-occurs when a schism opens between party leaders and traditional core-base voters on an issue of overriding national importance. Major parties survive periodic defeats, but they cannot survive disalignment.