The Boxtrolls dwell in the vast and intricate caverns under the cobblestone streets of Cheesebridge. Human legend has it the creatures are carnivorous monsters. In reality, though, they're just misunderstood oddballs who've raised an orphan human boy named Eggs. When Eggs discovers his true identity, he finds himself in the very centre of the ongoing struggle between his boxtroll family and cheese-loving fearmongerers. In the end, it takes a privileged little girl named Winnie to teach both Eggs and the people of Cheesebridge what it means to be human.